The Boss was born in  as the daughter of one of the original and higher-ranking members of the Wisemen's Committee , and later grew up in the care of the Philosophers. She learned from her father the most forbidden secrets of the Philosophers, and as a consequence, the shadowy organization arranged for his death. The Boss later went on to become an instructor at one of the Philosophers' "charm schools. She also planned the dummy raids on Heliopolis and the bombings of German air bases during the North African Campaign. On December 30, , she and Zero lost a close friend who had served under her in Layforce likely during Operation Crusader in Egypt. Zero delivered his SAS winged dagger pin to her after his death and she continued to keep it with her right up until her death. They participated in many special operations during World War II, contributing heavily to the Allied forces' victory. Likewise, she was also trained in sniping by The End.
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In the Metal Gear series, The Boss is a legendary American soldier, founder and leader of the Cobra unit, the biological mother of Ocelot ,  mentor and mother figure to Naked Snake , and is known as the mother of the U. Following Volgin's death, The Boss reveals she is the daughter of one of the original members of the Philosophers behind the Philosophers' Legacy. The whole mission is later revealed to be a coverup by the United States so that The Boss would steal Volgin's treasure and give it to Snake but then die at his hands to avoid a conflict between America and the Soviet Union. She was eventually commissioned into Project Mercury to test the Mercury capsule and launched into space at roughly the same time as Yuri Gagarin 's flight on April 12, However, the capsule suffered heavy damage upon reentry. The Boss, dubbed within the program as the "Mercury Lady", survived the crash but was rendered comatose for six months. All evidence of her role in the program was erased, including airbrushing her out of an official picture of the Mercury astronauts, she being the eighth member. The Boss's looks were inspired by actress Charlotte Rampling. The character was very well received by media. According to Eurogamer , "The Boss is often touted as being one of the strongest female characters in gaming.
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